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Vitamin C, which lemons are famously rich in, is a required co-factor for two specific enzymes: prolyl hydroxylase and lysyl hydroxylase

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Arnr ESJ, Sarioglu H, Lottspeich F, Holmgren A, Bck A (1999) High-level expression in Escherichia coli of selenocysteine-containing rat thioredoxin reductase utilizing gene fusions with engineered bacterial-type SECIS elements and co-expression with the sel A, sel B and sel C genes