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At the brush border membrane of enterocytes, an in-ward proton gradient is generated through the activity of an electroneutral proton/cation exchanger, Na + /H + antiporter

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During the following three decades, the hormonal status of glucagon was not generally accepted, but several groups had explored the possible mechanisms underlying the hyperglycaemia induced by glucagon (at that time it was also termed the hyperglycaemic-glycogenolytic factor due to its effect on glycogenolysis) [5]